compiled and edited by Kathryn Mauz, updated March 2021
hosted by the University of Arizona Herbarium (ARIZ), Tucson


INTRODUCTION John Reeder (1914-2009) and Charlotte Goodding Reeder (1916-2009) were agrostologists, or grass specialists, who devoted their careers to documenting the grass floras of North America and investigating topics in grass systematics and evolutionary biology. In retirement, "The Reeders," as they were known, were fixtures at the University of Arizona Herbarium for three decades. Read their story in Desert Plants vol.35(1): 25-42 (June 2019) here.
IN APPRECIATION For their years of scholarship and service, John and Charlotte were honored with a Lifetime Achievement Award at the first annual Arizona Botanists Meeting in 2003. At that time and after their passing in 2009, several colleagues offered stories of the Reeders' friendship and influence, featured under the In Appreciation tab.
PUBLICATIONS It is easy to imagine that Charlotte's interest in plants stemmed from her father's work and the frequent field trips her family made throughout the western regions where his investigations took place, and she evidently inherited the most fundamental qualifications of curiosity and intrepidness from her parents. As a plant collector, in addition to his official duties investigating problems in plant pathology and economic botany, Leslie Goodding undoubtedly passed the skills and habits of field botany along to his oldest daughter. Her own systematics investigations began in the grass genus Muhlenbergia. Charlotte's 1939 master's thesis led to the publication of five new species (the only species authored by "C.O.Goodd."), and she maintained her focus on the systematics of the grass family for the balance of her career. John also started down the botany path from an early age and, while his publications occasionally included treatments in other plant families, he became an agrostologist in every sense. John's investigations considered topics ranging from biogeography to gross morphology, anatomy, and cytology, with an overarching interest in grass systematics and evolutionary biology. They discovered many new species and added to the knowledge of regional grass floras, especially in the western United States and Mexico. In addition to sometimes illustrating grasses, Charlotte often assisted John in preparing papers for publication. The Reeders' publications are compiled under the Publications tab.
COLLECTIONS & TAXA Both Charlotte and John made floristic collections during their undergraduate and graduate days. Almost exclusively of grasses, the Reeders collected throughout Mexico from the 1950s–1970s, widely in the western states during John's tenure at the University of Wyoming, then further shifted their emphasis to the southwest after retiring to southern Arizona. Because of their tenure at several institutions during long careers, their wide-ranging collecting activities, and their diligence in making and distributing duplicate specimens, the Reeders' plant collections—with collection numbers exceeding 10,000—are housed among many herbaria in the Americas and elsewhere; notable among these are: the U.S. National Herbarium, the Yale Herbarium, the Rocky Mountain Herbarium, and the University of Arizona Herbarium. The Reeders' herbarium work involved not only critical study of their own material, but the annotation of numerous historical collections and the identification of thousands more specimens collected by others. Charlotte authored 8 new subgeneric taxa or new combinations, and published with John or others the names for 4 genera, 3 species, and 7 new combinations for subgeneric taxa. John named 2 suprageneric taxa and 34 new subgeneric taxa, and authored (some with other authors not including Charlotte) an additional 31 new combinations. A New Guinean fern was named in honor of John, its collector, and both John and Charlotte were honored by the names of three grass species and one grass genus, Reederochloa. These names are listed under the Taxonomy tab.

Primary herbaria holding specimens collected by Charlotte Goodding, or Charlotte Reeder, and John Reeder:

U.S. National Herbarium (US), Smithsonian Institution, Washington, D.C.

Oregon State University Herbarium (OSC), Corvallis

Yale Herbarium (YU), Peabody Museum of Natural History, New Haven, Ct.

Rocky Mountain Herbarium (RM), University of Wyoming, Laramie

University of Arizona Herbarium (ARIZ), Tucson

ARCHIVES The Reeders' botanical library was divided between the Deaver Herbarium at Northern Arizona University, in Flagstaff, and the University of Arizona Herbarium, in Tucson. A collection of their papers, notes, and photographs resides at the University of Arizona Libraries Special Collections. Their field notebooks are preserved at the University of Arizona Herbarium. An inventory of the collection is available under the Archives tab.


graphic adapted from an illustration of Muhlenbergia biloba by Charlotte Reeder, in Madroño 13: 249. 1956

Online supplement to Desert Plants, vol.35(1): 25-42, June 2019
compiled and edited by Kathryn Mauz, updated June 2019
hosted by the University of Arizona Herbarium (ARIZ), Tucson

Botanical publications by Charlotte Goodding or Charlotte Reeder, including those with John Reeder

Goodding, C.O. 1940. Two new species of Muhlenbergia. Journal of the Washington Academy of Sciences 30: 19-20.

Goodding, C.O. 1941. Three new species of Muhlenbergia. Journal of the Washington Academy of Sciences 31: 504-506.

Reeder, C.G. 1949. Muhlenbergia minutissima (Steud.) Swallen and its allies. Journal of the Washington Academy of Sciences 39: 363-367.

Reeder, C.G. 1956. Muhlenbergia brandegei, a new species from Baja California, Mexico, and its relationship to Muhlenbergia biloba. Madroño 13: 244-252.

Reeder, J.R. & C.G. Reeder. 1960. [in memoriam] Alexander William Evans (1868-1959). Rhodora 62: 245-250.

Reeder, J.R. & C.G. Reeder. 1963. Notes on Mexican grasses I: new and noteworthy species of Bouteloua. Brittonia 15: 215-221.

Reeder, J.R. & C.G. Reeder. 1963. Notes on Mexican grasses II: Cyclostachya, a new dioecious genus. Bulletin of the Torrey Botanical Club 90: 193-201.

Reeder, C.G. & J.R. Reeder. 1965. [book review] A Selected Guide to the Literature on the Flowering Plants of Mexico, by Ida Kaplan Langman. Bulletin of the Torrey Botanical Club 92: 494-500.

Reeder, J.R., C.G. Reeder & J. Rzedowski. 1965. Notes on Mexican grasses III: Buchlomimus, another dioecious genus. Brittonia 17: 26-34.

Reeder, J.R. & C.G. Reeder. 1966. Notes on Mexican grasses IV: dioecy in Bouteloua chondrosioides. Brittonia 18: 188-191.

Reeder, C.G. & J.R. Reeder. 1966. Notes on Mexican grasses V: two gypsophilous species of Muhlenbergia. Madroño 18: 185-192.

Reeder, J.R. & C.G. Reeder. 1968. Parodiella, a new genus of grasses from the high Andes. Boletín de la Sociedad Argentina de Botánica 12: 268-283.

Reeder, J.R. & C.G. Reeder. 1969. [meeting abstract] Cyclostachya stolonifera, a "rare" Mexican grass. Journal of the Colorado-Wyoming Academy of Science 6(2): 18-19.

Reeder, J.R. & C.G. Reeder. 1969. New name for Parodiella. Boletín de la Sociedad Argentina de Botánica 11(4): 239.

Reeder, J.R. & C.G. Reeder. 1969. Citología y distribución de Bouteloua karwinskii y B. chasei. Boletín de la Sociedad de Botánica de México 30: 113-120.

Reeder, J.R. & C.G. Reeder. 1970. A new combination in Trichoneura from the Galápagos Islands. Madroño 20: 253.

Wiggins, I.L. & D.M. Porter (eds.). 1971. Flora of the Galápagos Islands. Stanford: Stanford University Press.
     Reeder, J.R. & C.G. Reeder. Gramineae, pp.823-892 (with 6 illustrations by Charlotte Reeder).

Reeder, J.R. & C.G. Reeder. 1972. [meeting abstract] Cytotaxonomy of Buchloë dactyloides (Gramineae). Journal of the Colorado-Wyoming Academy of Science 7(2-3): 104-105.

Reeder, J.R. & C.G. Reeder. 1972. [book review] Intermountain Flora, Vascular plants of the Intermountain West, U.S.A. Vol. I: Geological and botanical history of the region, its plant geography and a glossary; The vascular cryptogams and the gymnosperms, by Arthur Cronquist, Arthur H. Holmgren, Noel H. Holmgren, and James L. Reveal. Bulletin of the Torrey Botanical Club 99: 148-149.

Reeder, J.R. & C.G. Reeder. 1972. [book review] A Revised Flora of Malaya: An illustrated systematic account of the Malayan flora, including commonly cultivated plants, Vol. III-Grasses of Malaya, by H.B. Gilliland and H.M. Burkill. Bioscience 22: 324.

Reeder, J.R. & C.G. Reeder. 1973. [meeting abstract] Blepharidachne, an anomalous eragrostoid grass. Journal of the Colorado-Wyoming Academy of Science 7(4): 29.

Reeder, J.R. & C.G. Reeder. 1974. [meeting abstract] Publication date of Fournier's Mexicanas Plantas, Pars II. American Journal of Botany 61(supplement): 49.

Reeder, C.G. & J.R. Reeder. 1974. Publication date of Fournier's Mexicanas Plantas, Pars II. Taxon 23: 543-547.

Reeder, J.R. & C.G. Reeder. 1975. [meeting abstract] The Mexican grass genus Chaboissaea. American Journal of Botany 62(supplement): 58-59.

Reeder, J.R. & C. Reeder. 1978. Tragus racemosus in Arizona. Madroño 25: 107-108.

Reeder, J.R. & C.G. Reeder. 1978. [book review] Intermountain Flora—Vascular Plants of the Intermountain West, U.S.A. Vol. 6: The Monocotyledons, by Arthur Cronquist, Arthur H. Holmgren, Noel H. Holmgren, James L. Reveal & Patricia K. Holmgren. Bulletin of the Torrey Botanical Club 105: 241-243.

Reeder, C.G. & J.R. Reeder. 1980. Rediscovery of Orcuttia fragilis (Gramineae). Phytologia 46: 341-343.

Reeder, J.R. & C.G. Reeder. 1980. Systematics of Bouteloua breviseta and B. ramosa (Gramineae). Systematic Botany 5: 312-321.

Reeder, C.G. & J.R. Reeder. 1980. [book review] Flora of Baja California, by Ira L. Wiggins. Bulletin of the Torrey Botanical Club 107: 553-556.

Gould, F.W. & R. Moran (eds.). 1981. The grasses of Baja California. San Diego Society of Natural History Memoir 12. San Diego: San Diego Society of Natural History.
     Reeder, C.G. Muhlenbergia, pp.67-78.

Reeder, C.G. & J.R. Reeder. 1981. [book review] A Flora of New Mexico, by William C. Martin & Charles R. Hutchins. Bulletin of the Torrey Botanical Club 108: 490-492.

Reeder, C.G. 1985. The genus Lycurus (Gramineae) in North America. Phytologia 57: 283-291.

Reeder, J.R. & C.G. Reeder. 1985. Notes on Arizona grasses. Desert Plants 7(1): 22-23.

Reeder, C.G. & J.R. Reeder. 1986. Agrostis elliottiana (Gramineae) new to Arizona and New Mexico. Phytologia 60: 453-458.

Reeder, C.G. & J.R. Reeder. 1988. Aneuploidy in the Muhlenbergia subbiflora complex (Gramineae). Phytologia 65: 155-157.

Reeder, J.R. & C.G. Reeder. 1988. Hilaria annua (Gramineae), a new species from Mexico. Madroño 35: 6-9.

Reeder, C.G. & J.R. Reeder. 1989. A further note on Agrostis elliotiana (Gramineae) in Arizona. Phytologia 67: 134-138.

Reeder, J.R. & C.G. Reeder. 1990. Bouteloua eludens: elusive indeed, but not rare. Desert Plants 10(1): 19-22, 31.

Reeder, C.G. & J.R. Reeder. 1991. [in memoriam] Jason Richard Swallen (1903-1991). Taxon 40: 697-698.

Davidse, G., M. Sousa Sánchez & A.O. Chater (eds.). 1994. Flora Mesoamericana, vol. 6: Alismataceae a Cyperaceae. Universidad Nacional Autónoma de México, Missouri Botanical Garden Press, and the Natural History Museum (London).
     Pohl, R.W., C.G. Reeder & G. Davidse. Sporobolus, pp.273-276. [online]
     Reeder, C.G. Muhlenbergia, pp.276-286. [online]

Reeder, C.G. & J.R. Reeder. 1995. The resurrection of a species: Muhlenbergia straminea (Gramineae). Phytologia 78: 417-427.

Martin, P.S., D. Yetman, M. Fishbein, P. Jenkins, T.R. Van Devender & R.K. Wilson (eds.). 1998. Gentry's Río Mayo plants: The tropical deciduous forest and environs of northwest Mexico. Tucson: University of Arizona Press.
     Reeder, J.R. & C.G. Reeder. Poaceae, pp.498-520.

Stevens, W.D., C. Ulloa Ulloa, A. Pool & O.M. Montiel (eds.). 2001. Flora de Nicaragua, vol. 3: Angiosperma. Missouri Botanical Garden Monographs in Systematic Botany 85. St. Louis: Missouri Botanical Garden Press.
     Reeder, C.G. Muhlenbergia, pp.2075-2078. [online]
     Pohl, R.W., C.G. Reeder & G. Davidse. Sporobolus, pp.2133-2136. [online]

Barkworth, M.E., K.M. Capels, S. Long & M.B. Piep (eds.). 2003. Flora of North America north of Mexico, vol. 25: Magnoliophyta: Commelinidae (in part): Poaceae, part 2. New York and London: Oxford University Press.
     Reeder, C. Lycurus, pp.200-203. [online]

Cartron, J.-L.E., G. Ceballos & R.S. Felger (eds.). 2005. Biodiversity, ecosystems, and conservation in northern Mexico. New York: Oxford University Press.
     Van Devender, T.R., J.R. Reeder, C.G. Reeder & A.L. Reina-G. Distribution and diversity of grasses in the Yécora region of the Sierra Madre Occidental of eastern Sonora, Mexico, pp.107-121.

Barkworth, M.E., K.M. Capels, S. Long, L.K. Anderton & M.B. Piep (eds.). 2007. Flora of North America north of Mexico, vol. 24: Magnoliophyta: Commelinidae (in part): Poaceae, part 1. New York and London: Oxford University Press.
     Additions, Corrections, and Comments (Flora of North America vol. 25): Spartina Schreb., p.791; Echinochloa P.Beauv., p.792.

Reeder, J.R. & C.G. Reeder. 2007. Two more alien grasses now at home in the continental U.S.A. Phytologia 89: 1-7.


Botanical publications by John Reeder, including with authors other than Charlotte

Reeder, J.R. 1943. The status of Distichlis dentata. Bulletin of the Torrey Botanical Club 70: 53-57.

Reeder, J.R. 1946. Notes on Papuasian Saxifragaceae. Journal of the Arnold Arboretum 27: 275-288.

Merrill, E.D. & J.R. Reeder. 1946. New plant names published by Amos Eaton between the years 1817 and 1840. Bartonia 24: 26-79.

Reeder, J.R. 1948. The Gramineae-Panicoideae of New Guinea. Journal of the Arnold Arboretum 29: 257-392 +pl. 1-7.

Reeder, J.R. 1950. New and noteworthy Graminiae from New Guinea. Journal of the Arnold Arboretum 31: 320-328.

Reeder, J.R. 1951. Setaria lutescens an untenable name. Rhodora 53: 27-30.

Reeder, J.R. 1951. A further note on Cercis canadensis in Connecticut. Rhodora 53: 246-247.

Reeder, J.R. 1951. A new species of Poa from Peru. Journal of the Washington Academy of Sciences 41: 295-296.

Reeder, J.R. & S.-Y. Cheo. 1951. Notes on Xanthoxylum & Fagara in China. Journal of the Arnold Arboretum 32: 67-72 +pl. 1-2.

Reeder, J.R. 1953. The embryo of Streptochaeta and its bearing on the homology of the coleoptile. American Journal of Botany 40: 77-80.

Reeder, J.R. 1953. Affinities of the grass genus Beckmannia Host. Bulletin of the Torrey Botanical Club 80: 187-196.

Reeder, J.R. & K. von Maltzahn. 1953. Taxonomic significance of root-hair development in the Gramineae. Proceedings of the National Academy of Sciences 39: 593-598.

Reeder, J.R. 1956. The embryo of Jouvea pilosa as further evidence for the foliar nature of the coleoptile. Bulletin of the Torrey Botanical Club 83: 1-4.

Reeder, J.R. 1956. Note on Echinochloa muricata. Rhodora 58: 331-332.

Row, H.C. & J.R. Reeder. 1957. Root-hair development as evidence of relationships among genera of Gramineae. American Journal of Botany 44: 596-601.

Reeder, J.R. 1957. The embryo in grass systematics. American Journal of Botany 44: 756-768.
     reprinted in R. Ornduff (ed.). 1967. Papers on Plant Systematics. Boston: Little, Brown and Company: pp.52-64.

Reeder, J.R. & M.A. Ellington. 1960. Calamovilfa, a misplaced genus of Gramineae. Brittonia 12: 71-77.

Reeder, J.R. 1960. The systematic position of the grass genus Anthephora. Transactions of the American Microscopical Society 79: 211-218.

Bailey, D.L. (ed.). 1961. Recent Advances in Botany, from lectures & symposia presented to the IX International Botanical Congress, Montreal, 1959, vol. I. [Canada]: University of Toronto Press.
     Reeder, J.R. The Natural Classification of the Gramineae (I.U.B.S. Symposium): Embryology: The grass embryo in systematics, pp.91-96.

Reeder, J.R. & H.F. Decker. 1961. [meeting abstract] Affinities of Stipa and Aristida. American Journal of Botany 48: 549.

Reeder, J.R. & K.J. Norstog. 1961. The status of Hierochloë nashii and its relationship to H. odorata. Bulletin of the Torrey Botanical Club 88: 77-84.

Reeder, J.R. 1962. The bambusoid embryo: a reappraisal. American Journal of Botany 49: 639-641.
     reprinted in R. Ornduff (ed.). 1967. Papers on Plant Systematics. Boston: Little, Brown and Company: pp.65-67.

Reeder, J.R. 1965. [meeting abstract] Dioecy in the genus Bouteloua (Gramineae). American Journal of Botany 52: 650.

Reeder, J.R. 1965. The tribe Orcuttieae and the subtribes of the Pappophoreae (Gramineae). Madroño 18: 18-28.

Reeder, J.R. 1966. [meeting abstract] Validity of the monotypic tribe Jouveeae (Gramineae). American Journal of Botany 53: 635.

Löve, Á. (ed.). 1966. IOPB Chromosome Number Reports VI: contributions by J.R. Reeder. Taxon 15: 117.

Reeder, J.R. 1967. [meeting abstract] Validity of the tribe Spartineae (Gramineae). American Journal of Botany 54: 656.

Reeder, J.R. 1967. Notes on Mexican grasses VI: miscellaneous chromosome numbers [1]. Bulletin of the Torrey Botanical Club 94: 1-17.

Löve, Á. (ed.). 1967. IOPB Chromosome Number Reports XI: contributions by J.R. Reeder and J.R. Reeder & D.N. Singh. Taxon 16: 215-216.

Reeder, J.R. & D.N. Singh. 1967. Chromosome number in Calamovilfa. Bulletin of the Torrey Botanical Club 94: 199-200.

Reeder, J.R. 1967. Notes on Mexican grasses VII: new name for a dioecious endemic. Brittonia 19: 244.

Reeder, J.R. 1968. [meeting abstract] Systematic position of the genus Triniochloa (Gramineae). American Journal of Botany 55: 735.

Löve, Á. (ed.). 1968. IOPB Chromosome Number Reports XVI: contributions by J.R. Reeder & T.R. Soderstrom. Taxon 17: 203-204.

Reeder, J.R. 1968. Notes on Mexican grasses VIII: miscellaneous chromosome numbers, 2. Bulletin of the Torrey Botanical Club 95: 69-86.

Reeder, J.R. & D.N. Singh. 1968. Chromosome numbers in the tribe Pappophoreae (Gramineae). Madroño 19: 183-187.

Löve, Á. (ed.). 1969. IOPB Chromosome Number Reports XXII: contributions by J.R. Reeder, T.R. Soderstrom & C.E. Calderón. Taxon 18: 441-442.

Reeder, J.R. 1969. Las gramineas dioicas de México. Boletín de la Sociedad de Botánica de México 90: 121-126.

Reeder, J.R. 1970. [meeting abstract] Cytotaxonomy of Blepharoneuron. Journal of the Colorado-Wyoming Academy of Science 7(1): 37.

Reeder, J.R. & D.J. Crawford. 1970. [meeting abstract] Affinities of E[r]ioneuron and Munroa (Gramineae). American Journal of Botany 57: 752.

Reeder, J.R. 1971. Notes on Mexican grasses IX: miscellaneous chromosome numbers, 3. Brittonia 23: 105-117.

Reeder, J.R. 1972 [1971]. [meeting abstract] Cytotaxonomy of Spartina (Gramineae). Journal of the Colorado-Wyoming Academy of Science 7(2-3): 25.

Reeder, J.R. 1975. Symposium: "Evolution of Biotic Communities—The Grassland Biota" (First International Congress of Systematic and Evolutionary Biology, Boulder, Colo., 1973). Taxon 24: 51.

Reeder, J.R. 1976. Systematic position of Redfieldia (Gramineae). Madroño 23: 434-438.

Reeder, J.R. 1977. Chromosome numbers in western grasses. American Journal of Botany 64: 102-110.

Reeder, J.R. 1977. The "germination flap" in certain Gramineae. Madroño 24: 123-124.

Hammel, B.E. & J.R. Reeder. 1979. The genus Crypsis (Gramineae) in the United States. Systematic Botany 4: 267-280.

Reeder, J.R. 1980. Nomenclatural changes in Orcuttia (Gramineae). Phytologia 47: 221.

Reeder, J.R. 1981. The type locality of Orcuttia fragilis. Taxon 30: 308.

Reeder, J.R. 1981. (580) Proposal to conserve Centotheca (Gramineae). Taxon 30: 348-349.

Reeder, J.R. 1982. Systematics of the tribe Orcuttieae (Gramineae) and the description of a new segregate genus, Tuctoria. American Journal of Botany 69: 1082-1095.

Löve, Á. (ed.). 1984. Chromosome Number Reports LXXXII: contributions by J.R. Reeder. Taxon 33: 132-133.

Reeder, J.R. 1984. A new record for Schizachyrium semitectum in Mexico. Phytologia 55: 252.

Reeder, J.R. 1985. [book review] Grasses: A Guide to Their Structure, Identification, Uses and Distribution in the British Isles, by C.E. Hubbard & J.C.E. Hubbard. Taxon 34: 745.

Reeder, J.R. 1986. Mistaken identity in annual Eragrostis (Gramineae). Phytologia 60: 95-97.

Reeder, J.R. 1986. Another look at Eragrostis tephrosanthos (Gramineae). Phytologia 60: 153-154.

Reeder, J.R. 1986. Type specimen of Bouteloua ramosa Scribn. ex Vasey (Gramineae). Taxon 35: 149-153.

Reeder, J.R. & L.J. Toolin. 1987. Scleropogon (Gramineae), a monotypic genus with disjunct distribution. Phytologia 62: 267-275.

Reeder, J.R. & L.J. Toolin. 1988. Pappophorum philippianum (Gramineae) new to North America. Phytologia 64: 402-403.

Reeder, J.R. & L.J. Toolin. 1989. Notes on Pappophorum (Gramineae: Pappophoreae). Systematic Botany 14: 349-358.

Reeder, J.R. & R.S. Felger. 1989. The Aristida californica-glabrata complex (Gramineae). Madroño 36: 187-197.

Reeder, J.R. 1991. A new species of Panicum (Gramineae) from Arizona. Phytologia 71: 300-303.

Reeder, J.R. 1993. What is Aristida peruviana? Madroño 40: 266-267.

Hickman, J.C. (ed.). 1993. The Jepson manual: higher plants of California. Berkeley: University of California Press.
     Reeder, J.R. Eragrostis, pp.1257-1258 [online]; Neostapfia [online], Orcuttia, pp.1276-1277 [online]; Tuctoria, p.1300 [online].

Reeder, J.R. 1994. Setaria villosissima (Gramineae) in Arizona: fact or fiction. Phytologia 77: 452-455.

Reeder, J.R. 1994. Stipa tenuissima (Gramineae) in Arizona—a comedy of errors. Madroño 41: 328-329.

Peterson, P.M. & J.J. Ortíz-Diaz. 1998. Allelic variation in the amphitropical disjunct Muhlenbergia torreyi (Poaceae: Muhlenbergiinae). Brittonia 50: 381-391.
     Reeder, J.R. Appendix I: Chromosome counts of Muhlenbergia arenicola and M. torreyi, p.388.

Toolin, L. & J.R. Reeder. 2000. The status of Setaria macrostachya and its relationship to S. vulpiseta (Gramineae). Systematic Botany 25: 26-32.

Reeder, J.R. 2001. Chromosome number in Choisya (Rutaceae). Crossosoma 26: 12.

Reeder, J.R. 2001. Noteworthy collections: Setaria arizonica Rominger, Setariopsis auriculata (E. Fourn.) Scribner, Alopecurus arundinaceus Poir., Tridens albescens (Vasey) Wooton & Standley, Urochloa panicoides P. Beauv., Sclerochloa dura (L.) P. Beauv., Enneapogon cenchorides (Licht.) C.E. Hubbard (Gramineae). Madroño 48: 211-213.

Barkworth, M.E., K.M. Capels, S. Long & M.B. Piep (eds.). 2003. Flora of North America north of Mexico, vol. 25: Magnoliophyta: Commelinidae (in part): Poaceae, part 2. New York and London: Oxford University Press.
     Reeder, J.R. Scleropogon, pp.42, 44, 46; Pogonarthria, pp.105-107 [online]; Pappophoreae, pp.285-289 [online]; Orcuttieae, pp.290-296 [online]; Setariopsis, pp.539-540 [online].
     Hammel, B.E. & J.R. Reeder. Crypsis, pp.139-141 [online].

Felger, R.S., T.L. Burgess, S. Dorsi, J.R. Reeder & T.R. Van Devender. 2005. Dichanthium (Poaceae) new to Arizona: open door for a potentially invasive species. Sida Contributions to Botany 21: 1905-1908.

Reeder, J.R. & K. Mauz. 2009. Panicum coloratum new for Arizona, and Echinochloa holciformis new for the United States. Phytologia 91: 347-352.

Mauz, K. & J.R. Reeder. 2009. Marsilea mollis (Marsileaceae) sporocarps and associated insect parasitism in southern Arizona. Western North American Naturalist 69: 382-387.

Baldwin, B.G., D.H. Goldman, D.J. Keil, R. Patterson, T.J. Rosatti & D.H. Wilken (eds.). 2012. The Jepson manual: vascular plants of California. Berkeley: University of California Press.
     Reeder, J.R. Eragrostis, pp.1448 & 1450; Neostapfia, Orcuttia, pp.1468-1469; Tuctoria, p.1498.


Online supplement to Desert Plants, vol.35(1): 25-42, June 2019
compiled and edited by Kathryn Mauz, updated June 2019
hosted by the University of Arizona Herbarium (ARIZ), Tucson


Links are provided for type specimens and to illustrations prepared by Charlotte Reeder.
Nomenclatural changes are included in brackets; accepted names are shown in bold.

New taxa and new combinations published by Charlotte Goodding or Charlotte Reeder, including with John Reeder

POACEAE Muhlenbergia appressa C.O.Goodd. J. Wash. Acad. Sci. 31: 504. 1941
  Muhlenbergia brevis C.O.Goodd. J. Wash. Acad. Sci. 31: 505. 1941
  Muhlenbergia dubioides C.O.Goodd.
   [=Muhlenbergia palmeri Vasey]
J. Wash. Acad. Sci. 30: 20. 1940
  Muhlenbergia pectinata C.O.Goodd. J. Wash. Acad. Sci. 31: 505-506. 1941
  Muhlenbergia xerophila C.O.Goodd.
   [=Muhlenbergia elongata Scribn. ex Beal]
J. Wash. Acad. Sci. 30: 19-20. 1940
  Bouteloua elata Reeder & C.Reeder Brittonia 15: 215 +figs. 1 & 2. 1963
  Buchlomimus Reeder, C.Reeder & Rzed.
   [=Bouteloua Lag.]
Brittonia 17: 29: 1965
  Buchlomimus nervatus (Swallen) Reeder, C.Reeder & Rzed.
   [≡Bouteloua nervata Swallen]

Brittonia 17: 30. 1965
  Chaboissaea decumbens (Swallen) Reeder & C.Reeder
   [=Muhlenbergia decumbens Swallen]
Phytologia 65: 156. 1988
  Chaboissaea subbiflora (Hitchc.) Reeder & C.Reeder
   [=Muhlenbergia subbiflora Hitchc.]
Phytologia 65: 156. 1988
  Cyclostachya Reeder & C.Reeder
   [=Bouteloua Lag. sect. Cyclostachya (Reeder & C.Reeder) P.M.Peterson, Romasch. & Y.Herrera]
Bull. Torrey Bot. Club 90: 195. 1963
  Cyclostachya stolonifera (Scribn.) Reeder & C.Reeder
   [≡Bouteloua stolonifera Scribn.]
Bull. Torrey Bot. Club 90: 196. 1963
  Hilaria annua Reeder & C.Reeder Madroño 35: 7. 1988
  Lorenzochloa Reeder & C.Reeder
   [nom. nov., ≡Parodiella Reeder & C.Reeder, non Parodiella Speg.; =Ortachne Nees ex Steud.]
Bol. Soc. Argent. Bot. 11(4): 239. 1969
  Lorenzochloa erectifolia (Swallen) Reeder & C.Reeder
   [≡Ortachne erectifolia (Swallen) Clayton]
Bol. Soc. Argent. Bot. 11(4): 239. 1969
  Lycurus setosus (Nutt.) C.Reeder
   [=Muhlenbergia alopecuroides (Griseb.) P.M.Peterson & J.T.Columbus]
Phytologia 57: 287. 1985
  Muhlenbergia brandegeei C.Reeder Madroño 13: 248-250 +fig. 1. 1956
  Muhlenbergia eludens C.Reeder J. Wash. Acad. Sci. 39: 365-366 +fig. 1b. 1949
  Muhlenbergia gypsophila C.Reeder & Reeder Madroño 18: 186-190 +fig. 1 a1-h1, fig. 2 a1-d1. 1966
  Parodiella Reeder & C.Reeder
   [=Ortachne Nees ex Steud.]
Bol. Soc. Argent. Bot. 12: 268. 1968
  Parodiella erectifolia (Swallen) Reeder & C.Reeder
   [≡Ortachne erectifolia (Swallen) Clayton]
Bol. Soc. Argent. Bot. 12: 279. 1968
  Trichoneura lindleyana Ekman var. albemarlensis (B.L.Rob. & Greenm.) Reeder & C.Reeder Madroño 20: 253. 1970

New taxa and new combinations published by John R. Reeder, including with authors other than Charlotte

ESCALLONIACEAE Polyosma amygdaloides Reeder J. Arnold Arbor. 27: 283. 1946
  Polyosma induta Reeder J. Arnold Arbor. 27: 284-285. 1946
  Polyosma mucronata Reeder J. Arnold Arbor. 27: 282-283. 1946
  Polyosma occulta Reeder J. Arnold Arbor. 27: 285-286. 1946
  Polyosma oligantha Reeder J. Arnold Arbor. 27: 286-287. 1946
  Polyosma vochysioides Reeder J. Arnold Arbor. 27: 285. 1946
PARACRYPHIACEAE Quintinia brassii Reeder J. Arnold Arbor. 27: 280 +fig. 1a-c. 1946
  Quintinia lanceolata Reeder J. Arnold Arbor. 27: 278-279. 1946
ROUSSEACEAE Carpodetus amplus Reeder J. Arnold Arbor. 27: 275-276. 1946
  Carpodetus archboldianus Reeder J. Arnold Arbor. 27: 276-277. 1946
  Carpodetus denticulatus (Ridl.) Reeder J. Arnold Arbor. 27: 277-278. 1946
  Carpodetus flexuosus (Ridl.) Reeder J. Arnold Arbor. 27: 278. 1946
  Carpodetus fuscus Reeder J. Arnold Arbor. 27: 277. 1946
  Carpodetus montanus (Ridl.) Reeder J. Arnold Arbor. 27: 278. 1946
RUTACEAE Fagara dissita (Hemsl.) Engl. var. hispida Reeder & S.Y.Cheo
   [≡Zanthoxylum dissitum Hemsl. var. hispidum (Reeder & S.Y.Cheo) C.C.Huang]
J. Arnold Arbor. 32: 69. 1951
  Fagara mollis (Rehder) Reeder & S.Y.Cheo
   [≡Zanthoxylum molle Rehder]
J. Arnold Arbor. 32: 69. 1951
  Fagara oxyphylla (Edgew.) Reeder & S.Y.Cheo
   [≡Zanthoxylum oxyphyllum Edgew.]
J. Arnold Arbor. 32: 69. 1951
  Fagara rhetsoides (Drake) Reeder & S.Y.Cheo
   [=Zanthoxylum myriacanthum Wall. ex Hook.f. var. myriacanthum]
J. Arnold Arbor. 32: 69. 1951
  Fagara robiginosa Reeder & S.Y.Cheo
   [≡Zanthoxylum robiginosum (Reeder & S.Y.Cheo) C.C.Huang]
J. Arnold Arbor. 32: 68-69 +pl. 1. 1951
  Zanthoxylum acanthopodium DC. var. deminutum (Rehder) Reeder & S.Y.Cheo J. Arnold Arbor. 32: 71. 1951
  Zanthoxylum arenosum Reeder & S.Y.Cheo J. Arnold Arbor. 32: 70 +pl. 2. 1951
  Zanthoxylum simulans Hance var. imperforatum (Franch.) Reeder & S.Y.Cheo
   [=Zanthoxylum bungeanum Maxim. var. bungeanum]
J. Arnold Arbor. 32: 70. 1951
POACEAE Andropogon brevifolius Sw. var. cryptopodus (Ohwi) Reeder J. Arnold Arbor. 29: 363. 1948
  Andropogon micranthus Kunth var. multicispiculus (Ohwi) Reeder J. Arnold Arbor. 29: 365-366. 1948
  Andropogon spiciger (S.T.Blake) Reeder
   [≡Capillipedium spicigerum S.T.Blake]
J. Arnold Arbor. 29: 366. 1948
  Anthoxanthum horsfieldii (Kunth ex Benn.) Mez ex Reeder J. Arnold Arbor. 31: 325-327. 1950
  Bouteloua eriostachya (Swallen) Reeder Bull. Torrey Bot. Club 94: 7. 1967
  Brachiaria coccosperma (Steud.) Stapf ex Reeder
   [=Urochloa villosa (Lam.) T.Q.Nguyen]
J. Arnold Arbor. 29: 273. 1948
  Brachiaria fusiformis Reeder
   [≡Urochloa fusiformis (Reeder) Veldkamp]
J. Arnold Arbor. 29: 274 +pl. 1. 1948
  Brachiaria subquadripara (Trin.) Hitchc. var. piligera (F.Muell. ex Benth.) Reeder
   [≡Urochloa piligera (F.Muell. ex Benth.) R.D.Webster]
J. Arnold Arbor. 29: 273. 1948
  Calamagrostis gunniana (Nees) Reeder J. Arnold Arbor. 31: 323. 1950
  Calamagrostis parviseta (Vickery) Reeder J. Arnold Arbor. 31: 324. 1950
  Calamagrostis pusilla Reeder
   [≡Deyeuxia pusilla (Reeder) Jansen]
J. Arnold Arbor. 31: 322. 1950
  Calamagrostis uncinoides (S.T.Blake) Reeder J. Arnold Arbor. 31: 324. 1950
  Chrysopogon filipes (Benth.) Reeder J. Arnold Arbor. 29: 360. 1948
  Chrysopogon filipes (Benth.) Reeder var. arundinaceus Reeder
   [=Chrysopogon filipes (Benth.) Reeder]
J. Arnold Arbor. 29: 360-361. 1948
  subtribe Cottinae Reeder Madroño 18: 25. 1965
  Cyrtococcum patens (L.) A.Camus var. warburgii (Mez.) Reeder
   [=Cyrtococcum patens (L.) A.Camus var. patens]
J. Arnold Arbor. 29: 286. 1948
  Digitaria abortiva Reeder
   [=Digitaria ciliaris (Retz.) Koeler]
J. Arnold Arbor. 29: 291 +pl. 2. 1948
  Dimeria ciliata Merr. var. heteromorpha Reeder J. Arnold Arbor. 29: 325-326. 1948
  Dimeria dipteros Reeder J. Arnold Arbor. 29: 324 +pl. 6. 1948
  Dimeria monostachya Reeder J. Arnold Arbor. 29: 324 +pl. 5a,b. 1948
  Eragrostis pectinacea Nees var. miserrima (E.Fourn.) Reeder Phytologia 60: 154. 1986
  Eremochloa ciliaris (L.) Merr. var. elata Reeder
   [=Eremochloa ciliaris (L.) Merr.]
J. Arnold Arbor. 29: 351. 1948
  Eufournia Reeder
   [nom. nov., ≡Calamochloa E.Fourn., non Calamochloë Rchb.; =Sohnsia Airy Shaw]
Brittonia 19: 244. 1967
  Eufournia filifolia (E.Fourn.) Reeder
   [≡Sohnsia filifolia (E. Fourn.) Airy Shaw]
Brittonia 19: 244. 1967
  Eulalia irritans (R.Br.) Kuntze var. egregia Reeder
   [≡Pseudopogonatherum egregium (Reeder) Jansen]
J. Arnold Arbor. 29: 336 +pl. 7d,e. 1948
  Hemarthria subulata Reeder J. Arnold Arbor. 29: 350 +pl. 5c-e. 1948
  Hierochloë longifolia Reeder
   [≡Anthoxanthum redolens (Vahl) P.Royen var. longifolium (Reeder) Y.Schouten]
J. Arnold Arbor. 31: 325-326. 1950
  Isachne obtecta Reeder J. Arnold Arbor. 29: 313 +pl. 4. 1948
  Isachne villosa (Hitchcock) Reeder J. Arnold Arbor. 29: 314. 1948
  Ischaemum littorale Reeder J. Arnold Arbor. 29: 345 +pl. 7a-c. 1948
  Microstegium ciliatum (Trin.) A.Camus var. laxum (Nees ex Steud.) Reeder J. Arnold Arbor. 29: 338. 1948
  Orcuttia viscida (Hoover) Reeder
   [=Orcuttia californica Vasey]
Phytologia 47: 221. 1980
  tribe Orcuttieae Reeder Madroño 18: 20. 1965
  Panicum creperum Reeder
   [nom. nov., ≡Hemigymnia fusca Ridl., non Panicum fuscum Sw.]
J. Arnold Arbor. 29: 267. 1948
  Panicum mindanaense Merr. var. pilosum Reeder J. Arnold Arbor. 29: 270. 1948
  Panicum mohavense Reeder Phytologia 71: 300-301 +fig. 1. 1991
  Poa pearsonii Reeder J. Wash. Acad. Sci. 41: 295 +fig. 1b. 1951
  Rottboellia rottboellioides (R.Br.) Reeder
   [=Rottboellia rottboellioides (R.Br.) Druce]
J. Arnold Arbor. 29: 354. 1948
  Schizachyrium semitectum (Swallen) Reeder Phytologia 55: 252. 1984
  Setaria montana Reeder
   [=Setaria parviflora (Poir.) Kerguélen]
J. Arnold Arbor. 29: 304-305 +pl. 3. 1948
  Themeda gigantea (Cav.) Hack. var. novoguineensis Reeder
   [≡Themeda novoguineensis (Reeder) Jansen]
J. Arnold Arbor. 29: 374. 1948
  Tuctoria Reeder Amer. J. Bot. 69: 1090. 1982
  Tuctoria fragilis (Swallen) Reeder Amer. J. Bot. 69: 1090. 1982
  Tuctoria greenei (Vasey) Reeder Amer. J. Bot. 69: 1091. 1982
  Tuctoria mucronata (Crampton) Reeder
   [=Tuctoria greenei (Vasey) Reeder]
Amer. J. Bot. 69: 1091. 1982

Plants named by other authors in honor of Charlotte Goodding Reeder and John R. Reeder

THELYPTERIDACEAE Cyclosorus reederi Copel.
   [=Sphaerostephanos acrostichoides (Desv.) Holttum]
Amer. Fern J. 43: 12. 1953
  Named for John R. Reeder, collector, Finschhafen, New Guinea, 1944.
POACEAE Axonopus reederi G.A.Black
   [=Axonopus poiophyllus Chase]
Advancing Frontiers Pl. Sci. 5: 45 +t. 2. 1963
  Named for J.R. & C.G. Reeder, collectors, Chiapas, Mexico, 1953.
  Bouteloua reederorum J.T.Columbus
   [nom. nov., ≡Atheropogon stolonifer E.Fourn., non B. stolonifera Scribn.]
Aliso 18: 64. 1999
  Named in honor of J.R. & C.G. Reeder, "who have contributed much to our understanding of Bouteloua, including the discovery of dioecism in two species."
  Muhlenbergia reederorum Soderstr. Contr. U.S. Natl. Herb. 34: 122-124. 1967
  Named for J.R. & C.G. Reeder, collectors, "because of their interest in Gramineae, especially of Mexico, and the genus Muhlenbergia."
  Reederochloa eludens Soderstr. & H.F.Decker
   [≡Distichlis eludens (Soderstr. & H.F.Decker) H.L.Bell & Columbus]
Brittonia 16: 335 +figs. 1-10. 1964
  "We take pleasure in naming the new grass, Reederochloa, in honor of Professor John R. Reeder of Yale University, under whom both authors wrote their dissertations, and Mrs. Charlotte Goodding Reeder, a student of Muhlenbergia, whose suggestion to visit the Durango locality led to the discovery of the new genus."

Additional new taxa described by other authors from collections by Charlotte Goodding Reeder and John R. Reeder

TECTARIACEAE Tectaria heracleifolia (Willd.) Underw. var. maxima C.V.Morton
   [=Tectaria heracleifolia (Willd.) Underw.]
Amer. Fern J. 56: 126-127. 1966
  type coll. near Pueblo Nuevo, Veracruz, Mexico, 1953 (J.R. Reeder & C.G. Reeder 1975)
POACEAE Axonopus mexicanus G.A.Black Advancing Frontiers Pl. Sci. 5: 144. 1963
  type coll. near Villa Union, Sinaloa, Mexico, 1953 (J.R. Reeder & C.G. Reeder 2445)
  Bromus thysanoglottis Soderstr. & Beaman Publ. Mus. Michigan State Univ., Biol. Ser. 3(5): 509. 1968
  type coll. near Buenos Aires, Durango, Mexico, 1960 (J.R. Reeder & C.G. Reeder 3348)
  Scleropogon longisetus Beetle
   [=Scleropogon brevifolius Phil.]
Phytologia 49: 42-43. 1981
  type coll. south of Saltillo, Coahuilla, Mexico, 1963 (J.R. Reeder & C.G. Reeder 3626)


compiled by Amy Rule & Linda Kennedy, updated March 2021
hosted by the University of Arizona Herbarium (ARIZ), Tucson

Materials in the John R. Reeder and Charlotte Goodding Reeder Archives

Reeder field notebook and a mounted grass sample collected during their research.
(University of Arizona Libraries Special Collections)

FIELD NOTEBOOKS The Reeders' field notebooks are held at the University of Arizona Herbarium. A summary of their scope and contents, containing data for their plant collections as well as notes from their botanical travels between 1941 and 1996, is available here. For more information, please contact the Herbarium.
PAPERS The Reeders' personal and professional papers are held at the University of Arizona Libraries Special Collections. Materials include research notes, illustrations, and manuscripts; records of herbarium loans and specimen determinations; photographs from field work; correspondence; academic records; and printed matter related to their work and that of Charlotte's father, Leslie Goodding.

The collection is inventoried and available for use at the Special Collections reading room, University of Arizona Main Library, Tucson: MS 710, John R. Reeder and Charlotte Goodding Reeder Papers.


presented to

1st Annual Arizona Botanists Meeting
February 8, 2003
Desert Botanical Garden, Phoenix


compiled for the occasion by Tom Van Devender, Tucson, 2003

Words of appreciation...

Grasses seemed like such a mystery that I never thought I could effectively deal with them. But that was to change. John and Charlotte patiently pointed out my mistakes in identifications and where the existing keys had problems. They demystified grasses for me – what a privilege to get a start from the world experts. They patiently pointed out subtleties, and are always available to help. In 1987 John and I went on a field trip to northern Sonora. In the mountains west of Cananea he scrambled up a rocky slope and re-discovered the long-lost Bouteloua eludens in Mexico. I will never forget how he whooped and yelled: "Eludens Hill," 27.0 km E of Cananea on Mex Hwy 2, 1524 m, rocky south-facing slope with numerous grasses and forbs. What a privilege that we have a world-class collection of grasses at ARIZ – due to the devotion of Charlotte and John Reeder. Just last week I saw John patiently showing a student how to identify some grasses. Their friendship and professionalism and enthusiasm have helped shape my career. —Richard Felger, Drylands Institute and ARIZ, Tucson

John and Charlotte Reeder are unique among botanists – and botanists are truly a unique lot! I'm sure many of their colleagues will attest to the numerous important contributions they have made to grass taxonomy. I'd like to comment on the many and long days I spent in their company at the University of Arizona Herbarium during my graduate student days working on Asclepias phylogeny and Río Mayo Flora. I couldn't have had better company on those long quiet weekends in the basement dungeon of Shantz. Richard Felger, Victor Steinmann, and Larry Toolin were often there, but John and Charlotte were ALWAYS there. Just when I was at wits' end defining character states of milkweed coronas, John would save the day by interrupting me with a question, or I guess it was really a demand: "Would you just look at this?!" He would lead me over to the microscope where he was working to show me a horrendous abomination – a completely inaccurate description of a ligule in a key! "How could anyone make such a stupid mistake?" Of course, I had to agree with John, even on those occasions when I couldn't make out the subtlety that had raised his ire. Charlotte would just goad him on, shaking her head from side to side exclaiming, "Some people!".
     Best of all, of course, was lunch time. This was not optional. When John and Charlotte were ready for lunch, so was I! Charlotte would make strong tea, which I think pretty much fueled the assembly of my Asclepias data set. After lunch came the obligatory fight over cookies. Charlotte: "Here, Mark, take two." Mark: "No, thanks, I'm stuffed. I suppose I could eat one". Charlotte: "Oh, you really must have two." Mark: "Ok, I think I'll have two." It was during these lunches that I learned so much of history of botany and especially botanists of the United States. John and Charlotte would reminisce about young professor days at Yale and wild collecting trips in Mexico. I seem to remember something about them waking up early one morning in their car and being face-to-face with a mule! I sure miss those days! —Mark Fishbein, Mississippi State University Herbarium (MISSA), Starkville

I was a MS student at the RM in 1968. C.L. Porter had retired the previous spring leaving me with the freedom to shift from a treatment of the Caryophyllaceae for his Flora of Wyoming to doing a floristic inventory of Philmont Scout Ranch in northern New Mexico. Upon my return in late August, I met the Reeders. As the sole grad student in systematics, I received a great deal of much needed attention from John: introduction to Stebbins, Clausen et al., the Code, chromosome counting, grass classification, critical thinking, writing, manuscript editing, classical printing, marksmanship; from Charlotte, mothering, education in botanical illustration, bibliography, grass morphology, Aven Nelson and his students; and from the two, the scoop on who's who in botany.
     It is truly scary to think of my fate without these two industrious, knowledgeable, principled individuals. They changed my life far more than the vast majority of other mentors have. I love them dearly. —Ronald L. Hartman, Rocky Mountain Herbarium (RM), Laramie

John and Charlotte Reeder have been dear and trusted friends and colleagues of mine, and my wife Patty, since I started hanging around the herbarium. We have had countless professional and personal interactions over the years. What they have given to the University of Arizona Herbarium in positively the best, and the best curated, collection of Gramineae (as John would ask that I call them) in the southwestern part of the country and neighboring Mexico. In the past ten years that I have worked in the herbarium, they have been the sole curators of the grasses. "The best collection" includes the organization and accurate identification of all our 37 cabinets of that family. They steadily produce a list of publications that I believe equals and surpasses any one of our resident researchers. A very few examples include discovery of new species, contributions to the Jepson Manual and Flora Meso-Americana, and the grasses of the Galapagos Islands. Many of their publications are devoted to untangling the trail of confused nomenclature that taxonomists seem compelled to leave behind them. Charlotte has already left us a legacy of the wonderful native genus, Muhlenbergia. Just look at a list of Arizona "Muhlies" and look for her name to arise time and time again as the author. Don't miss C.O. Goodding, as she is the daughter of the famous Arizona botanist, Leslie Goodding, and she was publishing before she married John, after which her name has been listed as C.O. Reeder, or C.G. Reeder. On top of all this they are more than happy to do public service work, sometimes calling what people give them worthless scraps, yet making every effort to correctly identify them. They identify grasses for us all, and we have come to depend on them, perhaps too much. They are a wonderful asset to ARIZ, and I very much value their presence, their abilities and dedication, and I especially value their friendship. —Phil Jenkins, ARIZ

John and Charlotte Reeder are my role models. They love their work so much that they do not consider it work. They have led well-rounded professional lives, combining field and lab work, teaching, mentoring, service. They are open-minded, forward-looking and non-judgmental. They judge people by their professional performance and the content of their character. They are ever young in all the ways that count. Besides that, they know grasses and I am in awe of people who know grasses! —Lucinda McDade, Botany Department, Academy of Natural Sciences, Philadelphia

I first met John and Charlotte Reeder while I was an undergraduate at the University of Arizona in 1992. At that time grasses were an enigmatic group to me. I had no understanding of the difference between a glume and a lemma and couldn't even reliably distinguish a grass from a sedge. The Reeders' love and excitement for the grasses greatly moved me, and it is undoubtedly because of them that I presently view this group with a special interest and appreciation. On many an occasion they would remind me "Don't forget to collect grasses." Now when I am in the field, instead of ignoring and trampling these fascinating creatures, I try to distinguish and collect as many different kinds as possible. Whether it is pressing or identifying a grass, I always do, and always will, fondly think of John and Charlotte. Through their guidance and inspiration, I can readily distinguish a Bromus from a Festuca, a Piptochaetium from a Stipa, and an Eragrostis from a Poa. Had it not been for them, I doubt that I would be able to make this claim.
     Of the Reeders' many outstanding qualities, I have always been most impressed with their attention to detail. Their collections are always beautifully prepared, and they frequently advocate the use of "PCs" (paper clips, as Charlotte would say), small pieces of durable folder paper that are placed at the bends of a specimen to guarantee its proper posture when drying. The Reeders even personally mount many grass collections, both theirs and of others, in order to ensure optimal placement on the sheet. Especially noteworthy and impressive is Charlotte's collection of index cards with notes about and detailed illustrations of hundreds (if not thousands!) of Muhlenbergia specimens.
     Due to the Reeders' lively conversation, I greatly enjoyed my weekend lunch breaks in the University of Arizona Herbarium office, and it was during those meals that I developed a friendship with and great respect for John and Charlotte. The discussions were always varied and frequently full of interesting anecdotes about past botanists that the Reeders knew during their work at Harvard, Yale, and the University of Wyoming. Although I am now more than a thousand kilometers away and unable to visit Tucson regularly, on many a Saturday or Sunday at lunchtime I find myself drifting to Arizona in my mind. I can see the Reeders in the University of Arizona Herbarium office with their sandwiches, cookies and tea. At those times I only wish that it were easier to drop in for a noontime chat.
     It is most appropriate that John and Charlotte are honored at the first meeting of Arizona botanists. Their lifelong dedication to the grasses has greatly enhanced our knowledge of this important group in both Arizona and throughout the world. Their devotion and enthusiasm set a high standard for which other botanists should strive. I feel privileged to have benefited from the tutelage and friendship of the Reeders. John and Charlotte, thanks for everything! —Victor Steinmann, Instituto de Ecología, Centro Regional del Bajío, Pátzcuaro, Michoacán

In the late 1970s, I had become particularly interested in the grasses. While working with Tom Van Devender, I was attempting to identify grass flowers and other fragments preserved in packrat middens. The Reeders were intrigued by the idea that such identifiable bits could be as much as 40,000 years old. Their interest led them to help in my identifying work. In time, they furthered my interest in, and knowledge of, the Gramineae, as well as taxonomy, nomenclature, and bibliography. Their role as mentors and friends has continued for nearly 25 years now, and I'm glad of it! —Larry Toolin, ARIZ

I began as a herpetologist. In graduate school at the University of Arizona, Paul Martin introduced me to paleoecology and the "secrets of the past" entombed in ancient packrat middens. I reconstructed the Ice Age history of the Sonoran Desert vegetation and climate using seeds, leaves, and twigs preserved in the middens. To interpret the midden assemblages, I had to learn to recognize desert plants, beginning with Charles Mason's course in Systematic Botany, and Martin's Paleoecology and Man. I learned to recognize plants by their "gestalt" mostly wandering in the desert. On occasion, I'd crack my Kearney & Peebles and tackle a key. I began collecting plants as vouchers for the midden studies. Art Phillips once told me that making good herbarium specimens was an art, convincing three-dimensional plants to look good on two-dimensional herbarium sheets. It was important because specialists appreciate working on nice material.
     As a field botanist, natural historian, I have always relied on and worked with a network of dedicated systematists, so in love with their groups that they are excited to see new specimens from interesting places. My "gestalt" approach worked well for recognizing most desert plants – except grasses, which all looked alike. John and Charlotte Reeder changed that. Because they were always in the University of Arizona Herbarium and SO interested in grasses, I began to collect for them on my various excursions. Twenty years later, I am surely, and unintentionally, one of the most prolific grass collectors in the Southwest and northwestern Mexico! My grass flora of the roadside rest areas of New Mexico and Texas is nearly complete! It was always a treat to find a "goodie" for the Reeders. It was usually something that appeared nondescript to me.
     In recent years, my conversion to floristics is mostly complete. The Reeders have identified the grasses from intensive surveys on the Río Cuchujaqui, Yécora area, and the la frontera border region in Sonora, and the Tucson Mountains and Ironwood Forest National Monument in Arizona. We are even coauthors on a new chapter on the distribution and diversity of the grasses of the Municipio de Yécora for a book on the biodiversity of northwestern Mexico!
     Visiting with the Reeders on weekends is always interesting. John identifies our specimens before the ink can dry in my catalog. He delights in showing me details of the glumes and florets under a microscope. He and Charlotte host weekend lunches in the Herbarium, pouring tea from a porcelain pot that is surely older than my house. This is a time to tell stories. John and Charlotte have had such a full life and done so many things – and remember them all. They are a story-telling team – Charlotte reminds John of a story to tell, and then fills in the gaps. And such stories – of botanists who I only know as authors of grass names (Merrill, Soderstrom, Aunt Agnes [Chase], etc.) or from their photographs on the Reeders' office wall, adventures in Mexico, of discovering new genera and species, stories about "Pappy" (pioneer Arizona botanist Leslie Goodding, Charlotte's father), etc.
     The Reeders' legacy here is enormous. The ARIZ grass collection contains all of their Mexican collections. They have personally identified or checked most of the specimens. John personally files the specimens. Specimens labeled "Determined J.R. & C.G. Reeder" in all Arizona, Sonora, and Baja California herbaria are wonderful resources in correctly identifying grasses and helping other botanists sort out the details in complex groups including Bouteloua (John's specialty) and Muhlenbergia (Charlotte's specialty). We are so fortunate that the Reeders "retired" to Arizona!
—Tom Van Devender, Arizona-Sonora Desert Museum, Tucson

It is impossible for me to think about the University of Arizona without thinking of the Reeders. They were a fixture when I showed up at the herbarium as a confused young undergraduate in 1978 and they remain so today. During those years, John and Charlotte have had a profound influence on my evolution as a botanist. John and Charlotte were instrumental in my early taxonomic training, but even more importantly, they placed the field into a national and international context. As I grew more interested in plant life, it was John and Charlotte who humanized the "legendary heroes of botany" with stories of their past interactions with these men and women, making them real people in my mind, folks who might one day be considered colleagues rather than just names authoring epithets in some recent revision. Stories of those who had ascended to the great herbarium in the sky, like Charlotte's "pappy", Leslie Goodding, made an especially big impression to this wide-eyed neophyte.
     It was due to John and Charlotte's influence that I was invited on my first major field trip to central and southern Mexico, with Alan Beetle in 1981. They also provided guidance during the years when my Master's research on Lennoaceae was beginning to shape my professional goals as a systematist. I always appreciated their invitations to come over to their work area, when they would press me for my impressions of some especially curious plant under the scope at the time.
     Since leaving Tucson for other places, I have counted on the Reeders for the latest news and gossip on the herbarium and its denizens. Our more or less annual dinner get-togethers have become a cherished tradition, and they are among my longest-standing friends and colleagues in botany. In fact, it is impossible for me to think about the U of A without thinking of the Reeders. —George Yatskievych, Missouri Botanical Garden (MO), St. Louis

compiled by Ben Wilder, Tucson, 2009

Words in memoriam...

I began my master's degree with C. L. Porter not realizing that he would be retiring the following Spring, 1968. Upon returning from a summer of floristics at Philmont Scout Ranch, I noted a stack of boxes in the RM. Shortly, the Reeders appeared. Henceforth, "Ronito" was treated as their son. John and Charlotte were both superb mentors and "parents" in so many ways.
     John introduced me to chromosomes. I would sit for hours next to him, listening as he cruised a slide and then treated me to a beautiful meiotic squash, a grass of course. All along he was instructing me on techniques and educating me about systematics, and life. Lunch of cheese and crackers was not to be missed with his reminiscences, talk of colleagues, and descriptions of travel through Mexico, and with frequent interjections of details by Charlotte. As his only graduate student, working on the caryophylls of Wyoming, of all things, I received their constant attention and guidance. They taught me to write, search the classics of systematics and evolution, and introduced me to the "masters" at the Seattle Congress in 1969.
     Upon receiving my graduate degree, I have had little contact with them. Yet, I am so happy to have been in their "intellectual and parental atmosphere". Despite the separation, the memories are vivid and treasured and I will always owe John and Charlotte for the solid academic foundation they provided. —Ronald L. Hartman, Rocky Mountain Herbarium (RM), Laramie

Among the many positives of my nearly 10 year residence in Tucson (and curatorship of ARIZ), friendship with John Reeder stands out as a superlative. He was always supportive, encouraging, helpful, and grateful (as of course is Charlotte). His support of younger scientists, his willingness to advise on projects, his immediate assumption of a collegial stance and relationship are traits I try to emulate. I also try daily to emulate his life long commitment to learning, to activity of the mind, to never give up despite as he is famous for having said "this aging gig is not for the timid!" I have a picture of John and Charlotte (at work and certainly on a Saturday or Sunday or holiday!) in their funny little office in the basement of Shantz on my office window sill as a write. I consider them to be my honorary grandparents (I don't think they would think that too presumtuous of me!). —Lucinda McDade, Rancho Santa Ana Botanic Garden, Claremont

Before I met John in about 1983, my gestalt for recognizing plants had not yet broadened enough to include grasses; they all looked alike! But he and Charlotte identified them so quickly and were SO interested in them, that I began collecting grasses. With John, I always felt like a student with the master as he showed me obscure, but very important, characters on my specimens under the microscope. Without words we began a partnership that lasted over 25 years and yielded thousands of grass specimens from the Southwest and many parts of Mexico. I never knew which grass would be something unusual to be tussled with like a dog toy for days or weeks until its place in the grass order was understood. Their knowledge and hard work not only enriched the ARIZ collection but all of the various floristic studies I was involved in. Eventually we coauthored a paper on the grass diversity of the Municipio de Yécora in the Sierra Madre Occidental in eastern Sonora.
     My most cherished memories of John and Charlotte are of Sunday lunches in the Herbarium with my wife Ana Lilia. As we shared tea from the old brown, yellow spotted, chipped teapot, cookies, and coyotas from Hermosillo, we loved to hear them tell stories like lovingly rehearsed duets about great botanists as real humans and long ago field trips to Mexico. They were always keen to hear about our trips and the grasses we found, and especially news about Richard F., Phil J., Kathy M., George Y., Victor S., Larry T., Ben W., and other botanical family members. John was so cute getting every last cookie crumb with an old paring knife, and blushing when lovely 7-year-old Arehli gave him a kiss. I miss him as a great botanist, an interesting man, and a friend. He probably has an easier time identifying those pesky Bromus and Tripsacum now. —Tom Van Devender, Arizona-Sonora Desert Museum, Tucson

I have had the great fortune of spending many weekends in the past several years working in the University of Arizona herbarium in the presence of John and Charlotte Reeder. Soon after I started working with Richard Felger, I was made aware that I needed to meet the Reeder's. I will forever be grateful for that link.
     John shared many wonderful stories over lunches in the herbarium. I loved hearing the story about his service in WWII in New Guinea when he would convey his true location to Charlotte, a fact censored by the Army, by mentioning that he was in the proximity of the Type collection of a certain species. Charlotte who was then at the US herbarium in Washington D.C. explained that she would go over the office of Agnes Chase, the foremost grass specialist of her time (who the Reeders referred to as Aunt Agnes), and they would determine John's location by referencing the appropriate botanical literature. But one of my favorite moments with John was when he shared with me the history of his botanists' pick.
     I was discussing with John the correct way to collect a grass specimen. He clearly told me that a "top-snatched" specimen is not worth much. When collecting a grass in the field there is a tendency to just reach down and pull the grass up from its top, however this often results in leaving the roots of the plant behind, and the associated critical information about whether it is an annual or perennial and/or any subterranean cleistogenes that may be present. So I asked John, "What do you use to get the roots of a grass?" This turned out to be a good question. He went on to share:
     "When the Japanese were sinking our boats in WWII the Army started offering welding classes for free. So in the early morning I would take a class for a couple hours and I learned to weld. Then I was told that I could get a job welding in Vancouver, WA." Charlotte added, "We really needed the money." John continued, "So I took the graveyard shift and they had scraps laying around for us to practice with, and I made myself a pick." He used the crossbars of the suspension of an old model T. "It has a point on one side and a chisel on the other. It has been with me ever since." That was in 1942. He told me that if he ever lost it he would be devastated. He came close a couple times in the field. But both times after restless nights with little sleep they set out in the early morning and found it where they had been the prior day.
     Several months later I went to get some plant presses with John from the back of his pickup truck. Opening the back of the camper was like opening a small window into his world. The truck bed was very organized with cardboard neatly cut to fit to all the odd angles of the floor. A bench was erected behind the wall that separated the passenger cabin from the bed, which was filled with neatly placed boxes. I grabbed the two presses, using these little knee pillows to ease my process. He said, "Don't take that one, that is my working press." Referring to a press on the side of the cab neatly secured to the car by a rope.
     I asked John if by chance he had the pick he used to uproot grasses. He gave me a coy look and gestured for me to follow him to the passenger cabin of the truck. He unlocked the door and sure enough between the pedals and the driver's seat on the floor was the revered tool! He grabbed it with immense pride, and it was a true beauty. Strong and looking like new, the tool is about 10 inches long. The hollow lightweight handle transitioned smoothly via an expert weld into the chisel/pick top. John became filled with excitement when he described using the pick to collect Sporobolus airoides (an extremely thick rooted grass), having a look of total satisfaction with the utility of his old friend. I asked him what he called the tool. "It is a botanist pick", he said. He placed the pick carefully back under the drivers seat, ready for the next collection.
     Time spent sharing events from my life and stories and pictures from my travels, reciprocated with interest, encouragement, and a gateway to a world and time that passed before I came into being has led to a friendship that bridges two centuries and fills me with joy and wonder. I consider myself amazingly fortunate to have known John and to continue to be able to interact with and learn from Charlotte. This connection is a true highlight of my formative years in Tucson. —Ben Wilder (ARIZ), Tucson

Dear old friend, you'll be missed. I wonder what it is like in that great Herbarium in the Sky. Give my regards to Mrs. Chase, Dr. Swallen, Tom Soderstrom, and the others. You'll be busy for a long time, following along behind your old chum, A.A.B., correcting all of his misdeterminations, or perhaps his brush with divinity will have provided him with somewhat clearer vision.
     John, you've battled mightily this last year against a mountain of obstacles and against all odds. Perhaps now it's time to treat yourself to some relaxation. Get in your pickup and take a week off to relive a few of those road-trips through Mexico and the Southwest. Reacquaint yourself with those enchanting "Boutelouers" and some old friends in the field. Having ascended to a higher plane, you'll have the benefit of hindsight to find all of those elusive fertile culms and rhizomes quickly and without any troubles.
     Those of us left behind will have some mighty big boots to fill. It is no fun to realize that henceforth we'll be keying all of our unknown grasses by ourselves, not nearly as quickly or as well as you would have. The meals, the chats, the stories, and the gossip will all be diminished without your participation. Far beyond that, your leaving has created a void in our hearts that will be hard to fill. As I said, you'll be missed. A lot.
     So, dear old friend, this isn't really good-bye. Your impact on our lives has made a permanent impression, and you'll not be forgotten. Have fun in that great Herbarium in the Sky, and save us all a seat next to the microscope at the Reeder table. We may be a little late, but sooner or later we'll be joining you.
     Your friend, —George Yatskievych, Missouri Botanical Garden (MO), St. Louis

Memoria written by Phil Jenkins (ARIZ) for the Flora of North America Newsletter:

John Reeder (1914-2009)

Charlotte Goodding Reeder (1916-2009)


Photos: Charlotte and John in the 1950s (courtesy of James Goodding and the University of Arizona Herbarium)
and the Reeders at ARIZ in 2003 (courtesy of Tom Van Devender & Ana Lilia Reina-Guerrero)